List of transitional forms

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A transitional form is an organism that has features intermediate of its ancestors and progeny. The term is most common in evolution to refer to organisms that show certain features (wings, feathers, gills and so on) partly in development. In theory, every fossil is a transitional form if it has descendants and each living creature is a transition between its parent and its offspring. However, evolution is about the features of populations rather than individuals; the transition at the species level can be too small in fossils; so the list below concentrates on broad transitional features and the genus or larger group.

Contents

[edit] Definitions

A fossil (and the organism it came from) is usually considered transitional with respect to just two groups. (If more than two are considered, then the definition of "transitional" is fuzzier.) Individual organisms (or individual species), considered by themselves, cannot be divided into "transitional" and "non-transitional", rather like how individual towns can be labeled "between" and "not between"; the term is relative. However, in cases where two groups are very populous or well-known, and all of the intermediate species have gone extinct, any intermediate organisms's transitional nature becomes noteworthy and noticeable, and may hence be called "transitional" all by itself. To return to the analogy of geography, the United States has a region called "the Midwest" or (somewhat derogatory) "flyover country". Of course every town and region in the US is in the middle and to the West of somewhere, and can be flown over; the reason the Midwest is singled out as a "transitional" region is that it lies between the more populous and city-strewn East and West Coasts.

While some of the fossilized species in this list may be direct descendants and direct ancestors of the forms on either side of the transition, most are in fact close relatives of the "true" ancestral species. This is because evolution produces a tree of life on which there are many paths "up", forward in time, but only one true path "down", through the various parent species. (A mother species can have many daughters, but each daughter has only one mother.) The number of fossils directly ancestral to, for example, modern birds decreases as one travels down the geologic column. Alongside each of these true bird ancestors are dozens of fossilized cousin bird species whose lines eventually died out altogether, but whose morphology provides clues about the nature of the ancestral species.

Because of all this, the term transitional form is often used to mean organisms that lie between two groups as cousins, not just as ancestors. For example, the echidna is not ancestral to modern mammals, but retains features of the reptile-like mammals which other mammals (the therians) lost, and is therefore transitional between ancient reptiles and therian mammals. A generally interchangeable term is intermediate form. Most iconic transitionals are the close descendants or cousins of a directly ancestral species whose fossils have not been found, or which never fossilized to begin with. (Fossilization is extremely rare.)

[edit] According to creationists

According to creationists, each and every one of the following fossils is either misinterpreted by scientists, a fraud manufactured by scientists, or does not exist at all. They are utterly wrong in this claim - so they keep claiming it anyway.

Creationists also define transitional forms in inherently absurd or vague ways that cannot be met by the evidence. For example, they may insist that each of a proposed transitional organism's parts (its legs, eyes, wings) should be "partially formed" instead of "fully formed", even if those parts have nothing to do with the transition. (For example, that an ape-human transitional form would have half-length limbs, or only one eye.) What "partially formed" means is not always clear.

Even more nebulously, it is argued that a true transitional would be an "incomplete creature", a fantasy that creationists will contrast with the known fossil record of "fully formed" or "fully functional" creatures. A similar idea is that "real" transitionals would be Frankenstein's monsters, stitched-together assemblages of the parts of the two species. (It is akin to asserting that evolution could not be real unless the Piltdown hoax was also real, or asserting that if teenagers were really between children and adults, they would have baby heads on adult bodies.) In short, if biologists believe the species's members were healthy enough to survive for more than a few days, then it can't be transitional, because it must have been "fully formed". The thinking here may be a product of biological essentialism — a thing is either A or B, and something "between" A and B is logically impossible or convoluted.

Some creationists propose that the common ancestor of two species would necessarily look like a blend of them, such as Kirk Cameron's infamous Crocoduck. This ignores that the evolutionary model involves traits arising without having to be present in the common ancestor. Otherwise, the world's earliest life forms would somehow look like combinations of every species alive today. Also, a given distant ancestor species with living progeny (for example, the reptilian ancestor of ducks and crocodiles) will also have many different-looking descendants, not just two. Evolutionary lines do not "run out" of divergences any more than the surname Smith would run out of acts of reproduction.

Another mistake/lie made by creationist is that a transitional fossil has to be the direct descendent of one species and the direct ancestor of another. What they do not understand is the true definition of a transitional form. A transitional form illustrates an evolutionary link, as it can have features of two species, but have no other species as descendants. For example, your mother would be a 'transitional form' between you and your grandmother, as she shares traits with both of you. However, if your mother had a sister, she would also be a "transitional form" between you and your grandmother, having traits from both of you.

For example, apes and humans split from a common ancestor seven million years ago and both lineages are still around.

For this reason the concept of "missing link" is a misleading one. A transitional form does not need to be a perfect halfway house directly linking one group of organisms to another. "It merely needs to record aspects of evolutionary change that occurred as one lineage split from another."[11]

It is also sometimes argued that if key transitional traits of an intermediate species (for example, the wing claws of Archeopteryx) have not completely disappeared today, the species is not transitional. (The ostrich has wing claws). Of course, biology is under no obligation to "clear the slate" and make it so that extinct species' traits are utterly foreign to us. To assume otherwise is to assume presentism, that there is something "special" about the modern day. Eventually, perhaps, ostrich claws will disappear -- will this make them "retroactively" transitional?

When all hope is lost, creationists call the intermediate a "mosaic species" that "doesn't prove anything", and point out that God can create whatever he wants.

The species listed here are all thought to be close relatives of species descending from the first forms in the list and ancestral to the last ones. Each list represents some branch on the tree of life, its members being the sub-branches and "twigs" extending from this branch.

[edit] Transitions in vertebrates before the Cenozoic

This is a fairly incomplete list. Please help us expand it!

[edit] Invertebrate to Vertebrate

  • Unnamed Upper (U.) Pre-Cambrian chordate -- First to bear a primitive notochord; archaetypical chordate.
  • Pikaia gracilens -- Middle (M.) Cambrian chordate with lancelet-like morphology.
  • Haikouella -- Lower (L.) Cambrian chordate, first to bear a skull; archaetypical craniate.
  • Haikouichthys -- L. Cambrian quasi-vertebrate, intermediate in developing a vertebral column; archaetypical vertebrate. [1]
  • Conodonts -- U. Cambrian to Triassic quasi-vertebrates with spinal cord; "bug-eyed lampreys".
  • Myllokunmingia -- L. Cambrian vertebrate with primitive spinal column; oldest true crown-group vertebrate.
  • Arandaspis -- L. Ordovician vertebrate, armoured jawless fish (ostracoderm), oldest known vertebrate with hard parts known from (mostly) complete fossils.[2]

[edit] Jawless Fish to Jawed Vertebrate

  • Birkenia -- Silurian primitive, jawless fish, a typical member of the Anaspida[3][4]
  • Cephalaspis -- Silurian armoured jawless fish, archaetypical member of the "Osteostraca," sister group to all jawed vertebrates.
  • Shuyu -- Silurian to Devonian, armoured jawless fish belonging to Galeaspida, related to Osteostraca. Internal cranial anatomy very similar to the anatomy seen in basal jawed vertebrates[5]. This similarity is directly implied with the translation of its name, "Dawn Fish," with the implication that it represents the "dawn of jawed vertebrates."

[edit] Acanthodian to shark[6]

  • Ptomacanthus -- sharklike fish, originally described as an acanthodian fish: brain anatomy demonstrates that it is an intermediate between acanthodians and sharks.
  • Cladoselache -- primitive/basal shark.
  • Tristychius -- another sharklike fish.
  • Ctenacanthus -- primitive/basal shark.
  • Paleospinax -- sharklike jaw, primitive teeth.
  • Spathobatis -- Ray-like fish.
  • Protospinax -- Ancestral to both sharks and skates.

[edit] Primitive jawed fish to bony fish[7]

  • Acanthodians -- superficially similar to early bony fishes, and some have been identified as being the ancestors of sharks.
  • Palaeoniscoids -- primitive bony fishes.
  • Canobius, Aeduella -- palaeoniscoids with more advanced jaws.
  • Parasemionotus -- combination of modern cheeks with more primitive features, like lungs
  • Oreochima -- first teleost fish
  • Leptolepids -- vaguely herring-like ancestors of modern teleost fish. Lung modified into swim bladder.
  • Amphistium and Heteronectes -- percomorphs that demonstrate the transition of the eye location of flatfishes.

[edit] Fish to amphibian[8]

  • Paleoniscoids — both ancestral to modern fish and land vertebrates
  • Osteolepis — modified limb bones, amphibian like skull and teeth
  • Kenichthys — shows the position of exhaling nostrils moving from front to fish to throat in tetrapods in its halfway point, in the teeth
  • Eusthenopteron, Sterropterygion — fin bones similarly structured to amphibian feet, but no toes yet, and still fishlike bodily proportions
  • Panderichthys, Elpistostege — tetrapod-like bodily proportions.
  • Obruchevichthys — fragmented skeleton with intermediate characteristics, possible first tetrapod.
  • Tiktaalik — a fish with developing legs. Also appearance of ribs and neck.
  • Acanthostega gunnari—famous intermediate fossil. most primitive fossil that is known to be a tetrapod
  • Ichthyostega — like Acanthostega, another fishlike amphibian
  • Hynerpeton — A little more advanced then Acanthostega and Ichtyostega
  • Labyrinthodonts — still many fishlike features, but tailfins have disappeared
  • Gars — Fish with vascularized swim bladders that can function as lungs
  • Lungfish and Birchirs — fish that have lungs

[edit] Primitive to modern amphibians[9]

  • Temnospondyls
  • Dendrerpeton acadianum
  • Archegosaurus decheni
  • Eryops megacephalus
  • Trematops
  • Amphibamus lyelli
  • Doleserpeton annectens
  • Triadobatrachus primitive frog
  • Vieraella
  • Karaurus primitive salamander

[edit] Amphibian to reptile[10]

  • Proterogyrinus
  • Limnoscelis
  • Tseajaia
  • Solenodonsaurus
  • Hylonomus
  • Paleothyris

[edit] Early reptile to turtle[11]

  • Captorhinus
  • Scutosaurus
  • Odontochelys Semitestacea -- partial formation of a turtle shell, showing how the hard underbelly, or plastron, formed first.[12]
  • Deltavjatia vjatkensis
  • Proganochelys

[edit] Early reptile to diapsid (dinosaurs and modern reptiles except for turtles)[13]

  • Hylonomus
  • Paleothyris
  • Petrolacosaurus
  • Araeoscelis
  • Apsisaurus
  • Claudiosaurus
  • Planocephalosaurus
  • Protorosaurus
  • Prolacerta
  • Proterosuchus
  • Hyperodapedon
  • Trilophosaurus

[edit] Reptile to mammal[14]

  • Paleothyris
  • Protoclepsydrops haplous
  • Clepsydrops
  • Archaeothyris
  • Varanops
  • Haptodus
  • Dimetrodon
  • Sphenacodon
  • Biarmosuchia
  • Procynosuchus
  • Dvinia
  • Thrinaxodon
  • Cynognathus
  • Diademodon
  • Probelesodon
  • Probainognathus
  • Exaeretodon
  • Oligokyphus
  • Kayentatherium
  • Pachygenelus
  • Diarthrognathus
  • Adelobasileus cromptoni
  • Sinoconodon
  • Kuehneotherium
  • Eozostrodon
  • Morganucodon -- a transition between "mammal-like reptiles" and "true mammals".
  • Haldanodon
  • Peramus
  • Endotherium
  • Kielantherium
  • Aegialodon
  • Steropodon galmani
  • Vincelestes neuquenianus
  • Pariadens kirklandi
  • Kennalestes
  • Asioryctes
  • Procerberus
  • Gypsonictops
  • Juramaia
  • Eomaia
  • Sinodelphys

[edit] Dinosaur to bird[15]

  • Allosaurus --A large theropod with a wishbone
  • Compsognathus --A small coeleurosaur with a wishbone
  • Epidendrosaurus
  • Epidexipteryx
  • Scandoriopteryx
  • Gigantoraptor
  • Gobivenator
  • Mei
  • Saurornithoides
  • Sinovenator
  • Buitreraptor
  • Pyroraptor
  • Unenlagia
  • Graciliraptor
  • Bambiraptor
  • Balaur
  • Tsaagan
  • Dromaeosaurus
  • Sinosauropteryx
  • Protarchaeopteryx
  • Caudipteryx
  • Velociraptor
  • Deinonychus
  • Utahraptor
  • Achillobator
  • Oviraptor
  • Sinovenator
  • Beipiaosaurus
  • Lisboasaurus
  • Sinornithosaurus
  • Microraptor -- a feathered bird with distinctly dinosaurian characteristics, such as its tail.
  • Xiaotingia -- slightly earlier than Archaeopteryx, slightly more like a dinosaur and less like a bird
  • Archaeopteryx -- the famous bird-with-teeth.
  • Anchiornis
  • Baptornis
  • Rahonavis
  • Confuciusornis
  • Sinornis
  • Iberomesornis
  • Theriznosaurus
  • Nothronychus
  • Citipati
  • Falcarius
  • Alxasaurus
  • Chirostenotes
  • Avimimus
  • Khaan
  • Incisivosaurus
  • Caenagnathus
  • Troodon
  • Byronosaurus
  • Ingenia
  • Hesperonychus
  • Conchoraptor
  • Patagopteryx
  • Ambiortus
  • Hesperornis
  • Apsaravis
  • Ichthyornis
  • Columba One of many typical modern birds

[edit] Transitional mammalian fossils

[edit] Primates

  • Purgatorius -- the earliest primate-like organism
  • Plesiadapis -- Mammal closely related to primates.
  • Carpolestes -- Mammal closely related to primates
  • Archicebus -- First euprimate, or something very similar to it.
  • Omomys -- Tarsier-like organism
  • Eosimias -- Basal anthropoid
  • Amphipithecus -- Another basal anthropoid
  • Apidium -- The first, primitive monkey.
  • Propliopithecus -- Primitive New World Monkey
  • Darwinius masillae -- a link between earlier primates and later ones.

[edit] Non-human primate to human

[edit] Cetaceans

  • Indohyus -- a vaguely chevrotain-like or raccoon-like aquatic artiodactyl ungulate with an inner ear identical to that of whales.
  • Ambulocetus-- an early whale that looks like a mammalian version of a crocodile
  • Pakicetus -- an early, semi-aquatic whale, a superficially wolf-like animal believed to be a direct ancestor of modern whales.
  • Rhodocetus -- An early whale with comparatively large hindlegs: not only represents a transition between semi-aquatic whales, like Ambulocetus, and obligately aquatic whales, like Basilosaurus.
  • Basilosaurus -- A large, elongated whale with vestigial hind flippers: transition from early marine whales (like Rhodocetus) to modern whales
  • Dorudon -- A small whale with vestigial hind flippers, close relative of Basilosaurus.

[edit] External links

[edit] Transitional plant fossils

  • Williamsonia -- an early flowering plant ("stem angiosperm")

[edit] See also

[edit] Footnotes

  1. http://adsabs.harvard.edu/abs/2003Natur.421..526S
  2. http://en.wikipedia.org/wiki/List_of_transitional_fossils
  3. http://www.britannica.com/EBchecked/topic/66626/Birkenia
  4. http://en.wikipedia.org/wiki/Birkenia
  5. http://www.nature.com/nature/journal/v476/n7360/full/nature10276.html
  6. [1]
  7. [2]
  8. [3]
  9. [4]
  10. [5]
  11. [6]
  12. Article in the journal Nature
  13. [7]
  14. [8]
  15. [9]
  16. [10]
  17. http://www.bbc.co.uk/sn/prehistoric_life/human/human_evolution/first_europeans1.shtml
  18. Homo heidelbergensis Click on the link, 'Evolutionary tree'.
  19. http://www.nytimes.com/2010/05/07/science/07neanderthal.html
  20. Was Neanderthal sex from the front or from the back?
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